Rediscovering the ‘Hulimaia’ (Moho angustipluma): an extinct bird from the rainforests of Hawai‘i

Have you ever heard of the Kioea (Moho angustipluma)? According to early American naturalists, the native people of Hawai‘i hadn’t either. Endemic to Hawai‘i Island, Moho angustipluma (formerly in the genus Chaetoptila) is a species of Hawaiian ‘honeyeater’ that went extinct by the 1860s, and until recently, has been widely recognized by the native name “Kioea.” The assumed name of this tropical bird has persisted in literature since the late nineteenth century with little information available to contest it. However, new research by Matthew R. Halley, Ph.D., Research Director and Curator of Birds at the Delaware Museum of Nature and Science, sheds new light on this mysterious species. In a new paper in Pacific Science (University of Hawaii Press), Halley proposes that its true native name has been obscured by time: “Hulimaia.”

Three artistic representations of Moho angustipluma: (left) unnumbered lithographic plate from Wilson and Evans
(1891), presumably based on the Cambridge specimen (UMZC 27/mel/6/a/1), captioned “F. W. Frohawk del. et lith. /
West Newman, imp.”; (center) unnumbered lithographic plate from Rothschild (1900), presumably based on the Rothschild
specimen (AMNH458995), captioned “J.G. Keulemans del. et lith. /Mintern Bros. imp.”; (right) Pl. 11 from Cassin (1858),
based on the holotype (USNM 15771), captioned “T. R. Peale del. / W. G. Newton Sc.”

Titian Ramsay Peale’s (1849) original description of M. angustipluma stated that this bird “frequent[ed] the woody districts … generally found about those trees which are in flower.” This implies that the bird inhabited forested areas on Hawai‘i Island and relied on nectar as its primary food source. Specimens of M. angustipluma reach lengths of 320 to 345 mm, which makes this species the largest known nectivorous passerine (perching) bird species on Hawai‘I, so the name Kioea (standing tall) would seem to be a fitting description—but see below.

Its brown plumage, streaked with white, did not possess the typical black and yellow colors typical of Moho species, although recent genetic evidence suggests that it is correctly placed in that genus (Zhao et al. 2026). Moreover, although native Hawaiians frequently used native bird feathers in cultural materials and artifacts, especially those of the Hawai‘i ‘Ō‘ō (Moho nobilis)and Hawai‘i Mamo (Drepanis pacifica)—both also extinct—the feathers of M. angustipluma are evidently absent from documented Hawaiian feather work. The species is also not represented in native Hawaiian literature, cultural practices, or bird taxonomies.

Three views of the Washington specimen (USNM15771), holotype of Moho angustipluma (Peale), its three extant
labels, and its entry in the USNM ledger. The oldest label (top right) was created after the specimen was cataloged at USNM
on 15 December 1859, after Cassin (1858), and is now detached from the specimen and stored in a separate binder. Reproduced
courtesy of the Division of Birds, National Museum of Natural History, Smithsonian Institution.

In literary accounts, scholars concluded that the native Hawaiians, who failed to recognize the name “Kioea” when asked, knew nothing about this native species. However, Halley has proposed that traditional knowledge of M. angustipluma was a casualty of colonization—like the species itself, which went extinct within a century after European contact. Furthermore, he suggests that the reason that locals on Hawai‘i Island were not familiar with the name “Kioea,” was simply that it was not the bird’s true native name.

Halley’s research paper, “Forgotten Knowledge of the Extinct Hawaiian ‘Hulimaia’ (Mohoidae),” uncovers overlooked primary sources which address these misconceptions. Having studied the Peale family for many years, Halley had long puzzled over the history of M. angustipluma. It was not until he was introduced to the writings of Zepherin “Kepelino” Kahō’āli’i Teauotalani, a native Hawaiian scholar, through the scholarship Gomes (2020), that he broke new ground. Teauotalani lived on Hawai‘i during the period when M. angustipluma is believed to have gone extinct, and Halley argues that his account of the “Hulimaia, the Bird of the Mountain Banana Groves” is likely an overlooked native description of the species, which contains otherwise unknown information about its behavior and ecology.

Digital scan of Teauotalani’s (1859) description of “Ta Hulimaia, manu o ta ea.” Reproduced courtesy of the
He Aupuni Palapala Project, Bishop Museum Library & Archives.

Teauotalani (1859) described the “Hulimaia” in an essay titled “Huli-Toa Manu Havaii” (A description of Hawaiian birds), which appeared in a local Hawaiian-language newspaper. The name used by Teauotalani is onomatopoeic—i.e., the name imitates the sound of the bird’s vocalizations—as he stated when “[the bird] cries, it says ‘hulinm-mmaiah, hulinm-mmaiah’ therefore, as ‘hulinm-mmaiah is its cry, it is named hulimaia.” According to Halley, no extant Hawaiian species possesses the combination of large body size, brown plumage, and nectivorous feeding habits described by Teauotalani (1859). However, these features are a good match for the morphology and presumed behavior of the extinct M. angustipluma.

Further research showed that the name “Kioea” was simply an error, which had been introduced to literature by Sanford Dole, a non-native Hawaiian scholar, in 1879. The name simply became entrenched in literature after decades of uncritical repetition. Ultimately, this led Halley to the conclusion that “Hulimaia” (Teauotalani 1859) was the true native name of M. angustipluma.

In a world where science has historically favored Western (print) literature over the oral transfer of knowledge, to rediscover and restore even a small piece of indigenous knowledge—and a forgotten native name—is a massive success. However, Halley’s hope for the future is a simple one. He hopes that the Hulimaia—despite its tragic extinction—will come to be seen as a symbol of cultural restoration.

“Knowledge is something that predates the printing press,” Halley says, “Knowledge is something that ultimately comes from experience, and the native people of Hawai‘i have a great deal of that.”

Furthermore, he says that by recognizing the name “Hulimaia,” and speaking it aloud in the forests of Hawai‘i, we can help to preserve the cultural memory of a lost species. This will serve as a much-needed reminder, that we cannot take the native birds that still exist for granted.

Literature cited

Gomes, N. J. 2020. Reclaiming native Hawaiian knowledge represented in bird taxonomies. Ethnobiol. Lett. 11: 30–43.

Halley, M. R. 2026. Forgotten knowledge of the extinct Hawaiian ‘Hulimaia’ (Mohoidae). Pacific Science 80: 141–156.

Peale, T. R. 1849. United States Exploring Expedition . . . Volume 8. Mammalia and ornithology. C. Sherman, Philadelphia.

Rothschild, W. 1900. The avifauna of Laysan and the neighboring islands; with a complete history to date of the birds of the Hawaiian possessions. Part III. R. H. Porter, London.

Teauotalani, Z. 1859. Hoiliili Havaii: He Mau Hana, Olelo, Manao, E Pili Ana I To Havaii Nei, Pepa 3: He Vahi Huli-Toa Manu Havaii. Pai-Palapala Katolika.

Zhao, M., Kimball, R. T., & Braun, E. L. 2026. The phylogenetic position of the extinct Hawaiian honeyeaters: overcoming the limitations of antique DNA. Zoological Scripta 55: 1–15. https://doi.org/10.1111/zsc.70047.

Delaware Museum of Nature and Science opens to the public May 23

After an extensive $10.8 million, 17-month renovation project, the Delaware Museum of Nature and Science opens to the public on Monday, May 23. The museum, formerly the Delaware Museum of Natural History, closed at the end of 2020 for the project. All of the exhibits – many in place since 1972 – were removed and the walls were taken down to the studs. Installation of the new exhibits has been ongoing since the end of 2021.

“We’ve completely shed that dusty, old museum perception. The Delaware Museum of Nature and Science is dynamic, engaging, interactive, relevant, and modern,” Executive Director Halsey Spruance said. “Our focus is on what we know about nature and science, why it matters to us, and what we can do to protect the environment. There’s a huge emphasis on how we are all connected and how our actions matter.”

Grand Opening Weekend Events:  

Friday, May 20, 10 a.m. Ribbon cutting and museum tours for invited guests and the media

Saturday, May 21 and Sunday, May 22, 9:30 a.m.-4:30 p.m. Member-Only Preview, sponsored by M&T Bank | Wilmington Trust. Includes special tours about the renovation process and new exhibits from staff members throughout the weekend. Open to DelMNS members and members of partnering museums: Delaware Art Museum, Hagley Museum & Library, Mt. Cuba, Winterthur Museum, Garden & Library, and Tyler Arboretum. Admission is free for members, pre-registration for timed tickets is requested.

Monday, May 23, 9:30 a.m.-4:30 p.m. Open to the public. Timed tickets will be available at delmns.org in early May.

Visit Information: The museum will be open seven days a week from 9:30 a.m.-4:30 p.m. Admission price is $12.95 for ages 3 and up, $3.95 for toddlers ages 1-2, and free for infants under 12 months. There is a $1 discount for tickets purchased in advance online. Admission is free for DelMNS members. For the first year, members of Delaware Art Museum, Hagley Museum & Library, Mt. Cuba, Winterthur Museum, Garden & Library, and Tyler Arboretum also receive free general admission.

New galleries include:

  • Regional Journey Gallery: Stroll across a giant floor map of the state and explore deciduous and mixed forests, the Bald Cypress Swamp, a saltmarsh, dunes, and the Delaware Bay. Designated areas such as the Fair Play Foundation Field Station and the DuPont Research Headquarters provide an opportunity to learn what is happening in our local region and beyond.
  • Alison K. Bradford Global Journey Gallery: A giant floor map of the world occupies the center of this gallery, surrounded by three land-based ecosystems, including a tropical rainforest, Arctic tundra and African savanna, along with three different ocean environments (shallow, mid-water, and deep). The ecosystems demonstrate nature’s diversity, the interdependency of life, and how humans play the biggest role in change.
  • Ellice & Rosa McDonald Foundation PaleoZone: Meet the creatures that lived in the Mid-Atlantic during the Cretaceous Period. Skeletons of the fearsome Dryptosaurus dinosaur, the flying “bat lizard” Nyctosaur, and the aquatic giant Mosasaur are joined by smaller Cretaceous specimens.

Other new exhibit spaces include the Tree of Life in the atrium with a visual interpretation of the Tree of Life, depicting the evolution of organisms over billions of years and the relationships between them in increasingly diverse branches. Adjacent to the Atrium, the Bill & Denise Spence Discovery Gallery offers rotating, hands-on exhibits. Opening exhibits include Delaware Mineralogical Society, First State Robotics and the University of Delaware, in addition to the museum’s Collections & Research Division.

New amenities include the Rest, Relax, Recharge café with prepackaged sandwiches, salads and snacks from Jamestown Catering, along with coffee, water, and other beverages. The Delaware Community Foundation Respite Room is a dedicated, calming space for visitors with sensory challenges and developmental disorders to take a break, as well as being a quiet and private option for nursing parents.

In addition to the new galleries and public spaces, additional parts of the project included lighting and sound systems, installation of a fire alarm and fire suppression system, new paving, HVAC system, refreshed meeting, event and temporary exhibit spaces, and renovated restrooms.

Delaware Museum of Nature and Science Project Partners

  • Strategic Planning           Schultz and Williams
  • Project Manager             Aegis Property Group
  • Exhibit Designer             Reich&Petch
  • General Contractor        Bancroft Construction
  • Architect                         JacobsWyper
  • Exhibit Fabricator           Kubik Maltbie

Media Contact:
Jennifer Acord, Director of Communications
jacord@delmns.org