Rediscovering the ‘Hulimaia’ (Moho angustipluma): an extinct bird from the rainforests of Hawai‘i

Have you ever heard of the Kioea (Moho angustipluma)? According to early American naturalists, the native people of Hawai‘i hadn’t either. Endemic to Hawai‘i Island, Moho angustipluma (formerly in the genus Chaetoptila) is a species of Hawaiian ‘honeyeater’ that went extinct by the 1860s, and until recently, has been widely recognized by the native name “Kioea.” The assumed name of this tropical bird has persisted in literature since the late nineteenth century with little information available to contest it. However, new research by Matthew R. Halley, Ph.D., Research Director and Curator of Birds at the Delaware Museum of Nature and Science, sheds new light on this mysterious species. In a new paper in Pacific Science (University of Hawaii Press), Halley proposes that its true native name has been obscured by time: “Hulimaia.”

Three artistic representations of Moho angustipluma: (left) unnumbered lithographic plate from Wilson and Evans
(1891), presumably based on the Cambridge specimen (UMZC 27/mel/6/a/1), captioned “F. W. Frohawk del. et lith. /
West Newman, imp.”; (center) unnumbered lithographic plate from Rothschild (1900), presumably based on the Rothschild
specimen (AMNH458995), captioned “J.G. Keulemans del. et lith. /Mintern Bros. imp.”; (right) Pl. 11 from Cassin (1858),
based on the holotype (USNM 15771), captioned “T. R. Peale del. / W. G. Newton Sc.”

Titian Ramsay Peale’s (1849) original description of M. angustipluma stated that this bird “frequent[ed] the woody districts … generally found about those trees which are in flower.” This implies that the bird inhabited forested areas on Hawai‘i Island and relied on nectar as its primary food source. Specimens of M. angustipluma reach lengths of 320 to 345 mm, which makes this species the largest known nectivorous passerine (perching) bird species on Hawai‘I, so the name Kioea (standing tall) would seem to be a fitting description—but see below.

Its brown plumage, streaked with white, did not possess the typical black and yellow colors typical of Moho species, although recent genetic evidence suggests that it is correctly placed in that genus (Zhao et al. 2026). Moreover, although native Hawaiians frequently used native bird feathers in cultural materials and artifacts, especially those of the Hawai‘i ‘Ō‘ō (Moho nobilis)and Hawai‘i Mamo (Drepanis pacifica)—both also extinct—the feathers of M. angustipluma are evidently absent from documented Hawaiian feather work. The species is also not represented in native Hawaiian literature, cultural practices, or bird taxonomies.

Three views of the Washington specimen (USNM15771), holotype of Moho angustipluma (Peale), its three extant
labels, and its entry in the USNM ledger. The oldest label (top right) was created after the specimen was cataloged at USNM
on 15 December 1859, after Cassin (1858), and is now detached from the specimen and stored in a separate binder. Reproduced
courtesy of the Division of Birds, National Museum of Natural History, Smithsonian Institution.

In literary accounts, scholars concluded that the native Hawaiians, who failed to recognize the name “Kioea” when asked, knew nothing about this native species. However, Halley has proposed that traditional knowledge of M. angustipluma was a casualty of colonization—like the species itself, which went extinct within a century after European contact. Furthermore, he suggests that the reason that locals on Hawai‘i Island were not familiar with the name “Kioea,” was simply that it was not the bird’s true native name.

Halley’s research paper, “Forgotten Knowledge of the Extinct Hawaiian ‘Hulimaia’ (Mohoidae),” uncovers overlooked primary sources which address these misconceptions. Having studied the Peale family for many years, Halley had long puzzled over the history of M. angustipluma. It was not until he was introduced to the writings of Zepherin “Kepelino” Kahō’āli’i Teauotalani, a native Hawaiian scholar, through the scholarship Gomes (2020), that he broke new ground. Teauotalani lived on Hawai‘i during the period when M. angustipluma is believed to have gone extinct, and Halley argues that his account of the “Hulimaia, the Bird of the Mountain Banana Groves” is likely an overlooked native description of the species, which contains otherwise unknown information about its behavior and ecology.

Digital scan of Teauotalani’s (1859) description of “Ta Hulimaia, manu o ta ea.” Reproduced courtesy of the
He Aupuni Palapala Project, Bishop Museum Library & Archives.

Teauotalani (1859) described the “Hulimaia” in an essay titled “Huli-Toa Manu Havaii” (A description of Hawaiian birds), which appeared in a local Hawaiian-language newspaper. The name used by Teauotalani is onomatopoeic—i.e., the name imitates the sound of the bird’s vocalizations—as he stated when “[the bird] cries, it says ‘hulinm-mmaiah, hulinm-mmaiah’ therefore, as ‘hulinm-mmaiah is its cry, it is named hulimaia.” According to Halley, no extant Hawaiian species possesses the combination of large body size, brown plumage, and nectivorous feeding habits described by Teauotalani (1859). However, these features are a good match for the morphology and presumed behavior of the extinct M. angustipluma.

Further research showed that the name “Kioea” was simply an error, which had been introduced to literature by Sanford Dole, a non-native Hawaiian scholar, in 1879. The name simply became entrenched in literature after decades of uncritical repetition. Ultimately, this led Halley to the conclusion that “Hulimaia” (Teauotalani 1859) was the true native name of M. angustipluma.

In a world where science has historically favored Western (print) literature over the oral transfer of knowledge, to rediscover and restore even a small piece of indigenous knowledge—and a forgotten native name—is a massive success. However, Halley’s hope for the future is a simple one. He hopes that the Hulimaia—despite its tragic extinction—will come to be seen as a symbol of cultural restoration.

“Knowledge is something that predates the printing press,” Halley says, “Knowledge is something that ultimately comes from experience, and the native people of Hawai‘i have a great deal of that.”

Furthermore, he says that by recognizing the name “Hulimaia,” and speaking it aloud in the forests of Hawai‘i, we can help to preserve the cultural memory of a lost species. This will serve as a much-needed reminder, that we cannot take the native birds that still exist for granted.

Literature cited

Gomes, N. J. 2020. Reclaiming native Hawaiian knowledge represented in bird taxonomies. Ethnobiol. Lett. 11: 30–43.

Halley, M. R. 2026. Forgotten knowledge of the extinct Hawaiian ‘Hulimaia’ (Mohoidae). Pacific Science 80: 141–156.

Peale, T. R. 1849. United States Exploring Expedition . . . Volume 8. Mammalia and ornithology. C. Sherman, Philadelphia.

Rothschild, W. 1900. The avifauna of Laysan and the neighboring islands; with a complete history to date of the birds of the Hawaiian possessions. Part III. R. H. Porter, London.

Teauotalani, Z. 1859. Hoiliili Havaii: He Mau Hana, Olelo, Manao, E Pili Ana I To Havaii Nei, Pepa 3: He Vahi Huli-Toa Manu Havaii. Pai-Palapala Katolika.

Zhao, M., Kimball, R. T., & Braun, E. L. 2026. The phylogenetic position of the extinct Hawaiian honeyeaters: overcoming the limitations of antique DNA. Zoological Scripta 55: 1–15. https://doi.org/10.1111/zsc.70047.

New Bird Species Discovered in Collections: A Tawi-Tawi Tale of Mismatched Specimens

The Delaware Museum of Nature and Science (DelMNS) houses an expansive bird collection in its Collections & Research Division, with over 116,000 bird specimens and 37,000 egg clutches, which are used for both scientific research and education. This collection is managed by DelMNS Research Director and Curator Dr. Matthew Halley, who has worked full-time at the museum since June 2021.

Last year, Dr. Halley encountered something that made him take pause: a single white-throated specimen in a drawer of brown-throated Pink-bellied Imperial Pigeon specimens (Ducula poliocephala) from the Philippines. The peculiar white-throated specimen was collected on Tawi-Tawi Island in 1971 by Dioscoro S. Rabor.

In a paper published in the Bulletin of the British Ornithologists’ Club, Dr. Halley has designated this specimen (DMNH 14999) as the “holotype” of a new species—the Tawi-Tawi Imperial Pigeon (Ducula tawitawi)—the first new (living) species of imperial pigeon to be described to science since 1928.

Species in the genus Ducula (family Columbidae) are commonly known as “imperial pigeons.” They live in the rainforest canopy throughout the Indo-Pacific region, and can reach considerable sizes (35–45 cm). Because their diet primarily consists of fruit, they likely play an important role as seed dispersers in tropical forests.

On the left, three views of DMNH 14999 and (right) its label and entry in the Delaware Museum of Nature and Science, Wilmington, ledger; the specimen was identified (apparently by duPont, see text) as ‘Ducula poliocephala nobilis’ when it was catalogued in 1972 (Matthew R. Halley)

The last living Ducula species was described in 1928, nearly a century ago, and before that, it was 1896. For decades, it has been assumed that the scientific inventory of the world’s Ducula species was complete—until now. Unfortunately, around 20% of the ~353 species of pigeons and doves are threatened with extinction, and we can probably add D. tawitawi to the list. It is only found in the threatened rainforests of Tawi-Tawi, a single oceanic island.

Dr. Halley searched through museum databases to locate other Imperial Pigeon specimens from Tawi-Tawi, and contacted colleagues at the Philippine National Museum in Manila, but he only managed to locate two others—both in the Smithsonian National Museum of Natural History (USNM). They were collected during the Menage Expedition in 1891 by Dean Conant Worcester and Frank S. Bourns. Dr. Halley went to the USNM earlier this year, to examine these specimens, and found that they too had white throats like the DMNH specimen!

Halley also found evidence that the songs of D. tawitawi are unique. The mating songs of the closely related Pink-bellied Imperial Pigeon (brown-throated D. poliocephala) and White-bellied Imperial Pigeon (D. forsteni) consist of two “coos,” but recordings from Tawi-Tawi made by Desmond Allen, a British ornithologist and field guide author, consisted of 5–8 coos. Because songs are innate (genetically encoded) in pigeons and doves, this implies that these populations are reproductively isolated—more evidence that D. tawitawi was an undescribed species.

Geographic variation in throat colour in the Pink-bellied Imperial Pigeon Ducula [poliocephala]
complex, as demonstrated by adult males held in the Delaware Museum of Nature and Science, Wilmington:
(A) DMNH 4501: Samar, May 1970; (B) DMNH 66314: Sibuyan, June 1973; (C) DMNH 1701: Mindanao, June
1961; (D) DMNH 14999: Tawi-Tawi, December 1971; (E) DMNH 10576: Mindoro, March 1971; (F) DMNH
66318: Marinduque, April 1972; (G) DMNH 3501: Luzon, March 1969 (Matthew R. Halley)

Amazingly, this isn’t the first time Dr. Halley has uncovered a new kind of bird. In previous years, he led a team that described a cryptic new species (Catharus arcanus) and two new subspecies (C. opertaneus tenebris, C. berlepschi nebulus) of nightingale-thrush from Panama, Ecuador and Peru, respectively, and he has also described two new subspecies of tailorbirds (genus Orthotomus) from the Philippines. The holotypes of the tailorbird subspecies are in the DelMNS collection.

D. tawitawi is the first new Ducula species to be identified in almost one hundred years—without a doubt, there’s always something extraordinary to find in the collections at the Delaware Museum of Nature and Science!

References

Halley, M. R. 2022. A new subspecies of Black-headed Tailorbird Orthotomus nigriceps (Cisticolidae) and clarification of age-related plumage sequences. Journal of Asian Ornithology 38: 129–134

Halley, M. R., Catanach, T. A., Klicka, J., and J. D. Weckstein. 2023. Integrative taxonomy reveals hidden diversity in the Catharus fuscater (Passeriformes: Turdidae) complex in Central and South America. Zoological Journal of the Linnean Society 199: 228–262.

Halley, M. R. 2025. Plumage variation in White-eared Tailorbird Orthotomus cinereiceps (Cisticolidae) with a description of a new subspecies from Mindanao, Philippines. Journal of Asian Ornithology 41: 51–57.

Halley, M. R. 2026. A new species of imperial pigeon (Columbidae: Ducula) from Tawi-Tawi, Philippines. Bulletin of the British Ornithologists’ Club 146: 295–310.

Addressing a “Sea-rious” Situation: Julia Wingate’s Take on Why Beach Restoration Matters.

Making Waves for Change

Julia Wingate, a rising high school senior studying natural resources, has a very particular issue that she’s passionate about: beach restoration in Delaware. As a Work-Based Learning Intern from the Appoquinimink School District, Julia has worked with the Delaware Museum of Nature and Science to develop a Gallery Interpretation kit focusing on Delaware Beach Ecosystems, stressing the importance of maintaining these beaches’ qualities and explaining how their condition affects the lives of the organisms who reside in those environments.

Designed to help guests better understand the organisms that live, nest, and grow on Delaware’s beaches, Julia’s beach ecology kit addresses many things: the necessity of beach restoration for animal protection and environmental conservation, what exactly beach restoration is, how to understand animal habitats on Delaware beaches, and how to contribute to beach restoration yourself.

“Beach restoration has been happening for many years now,” Julia says, describing the local history behind the practice. “Lots of money has been spent on it, and yet not much progress has been made.”

This lack of visible change has further motivated Julia to support this cause and increase its visibility among the general public. Julia makes a point to emphasize that restoring Delaware’s beaches isn’t solely for human enjoyment, but rather for the purpose of sustaining the organisms that reside there. A multitude of creatures, such as the marine horseshoe crabs and the migratory red knots, depend on Delaware’s shores for nesting and nourishment.

Without considerable restorative efforts, Delaware’s beaches will become unfit to provide for these species and many others, which would in turn lead to greater environmental collapse. Therefore, the only “shore” solution is exactly what Julia so strongly advocates for; restoring Delaware’s beaches.

What is Beach Restoration?

According to Julia Wingate, beach restoration can be described in many ways. Straightforwardly, it’s an earnest attempt to preserve beaches for the benefit of both humans and animals. It can also be restoring beaches to their natural condition prior to human intervention. But ultimately, Julia describes it as a process that creates a clean and safe space for all, regardless of species.

A plethora of efforts take place throughout the process of beach restoration, some of which look very different from one another. For instance, one particular task may be planting vegetation like grass and shrubs to prevent flooding and erosion. Other efforts may be seen through tracking and monitoring migrating shorebirds, which keeps local government and other relevant organizations informed about migration patterns and the current population numbers of different species. And, of course, the most natural thing that comes to mind is pumping new sand on to Delaware’s beaches, an effort that visibly replenishes depleted beach systems by transferring new sand on to eroded coastal areas.

Ultimately, beach restoration is a multifaceted process that touches upon the many components that make up an ecosystem, affecting abiotic and biotic resources alike.

The Ins and Outs of Julia’s Gallery Interpretation Kit

In the new Gallery Interpretation Kit that Julia has developed, a diverse range of materials have been incorporated in order to improve the public’s understanding of beach ecology. This ranges from examples of the different organisms that reside there, to the different plants that help to reinforce the longevity of Delaware’s sand dunes. Here are just a few of the several components she’s included in her educational project:

Horseshoe Crab

Using a horseshoe crab model, Julia aims to improve the public’s familiarity with the aquatic organism. These creatures are abundant in Delaware, serving as a food source for seabirds and also holding an important role in the biomedical industry due to the beneficial properties of their blood.

As a result of erosion and other environmental factors accelerated by human activity, many horseshoe crab nesting grounds are destroyed. This significantly interferes with the horseshoe crabs’ life cycle, and also often leaves them trapped on beaches. Other issues like littering also pose a risk to these creatures, as trash may be mistakenly ingested, which incorporates microplastics into the food chain.

Rufa Red Knot Migration Pattern

The rufa red knot carries out an incredibly long migration, wintering at the tip of South America before continuing its journey upward to Northern Canada to breed. Along this lengthy journey is a brief respite in the Delaware Bay, where red knots rest and replenish themselves before migrating further north. Here in the bay, red knots depend on horseshoe crab eggs as a primary food source.

However, because erosion disrupts horseshoe crab nesting areas, red knots are threatened by a potential loss of food resources. Additionally, the issue of microplastic contamination in horseshoe crab eggs also poses a threat to red knots, making them inedible and harmful to consume. Therefore, it’s important that beach restoration becomes a priority in order to preserve these interconnected species.

Beach Grass

Beach grass, along with other plants such as northern bayberry, serves as an integral defense against issues like coastal erosion and storm-related flooding. Because of its deep and fibrous root structure, beach grass is able to act as a sand stabilizer, allowing dunes to accumulate and grow in size.

However, these plants can be easily disrupted, especially in their early stages of development. Julia notes, “It is important that [beachgoers] can recognize these plants and are made aware not to pick up, walk on, or disturb them when on the beach.”

Why Does This Matter?

Just by looking at these three examples alone, it is easy to see how each of these different organisms make up an integral part of Delaware’s beach ecosystems. These organisms possess deeply interconnected relationships, whether you observe their relationships through the lens of the food web, or through their relationship with coastal erosion. Ultimately, it’s important to remember that each of these components are equally integral to sustaining beach ecosystems, and beach restoration is an essential part of guaranteeing the longevity of these organisms and their environment.

What Can You Do to Help?

As a part of her Gallery Interpretation kit, Julia provides the names of several groups with whom you can volunteer and make a difference, consisting of both government and independent organizations. If you feel moved by Julia’s cause, consider supporting or volunteering with the groups below:

DNREC

Delaware’s Department of Natural Resources and Environmental Control (DNREC) offers a multitude of volunteering opportunities related to beach restoration, consisting of research opportunities, stewardship opportunities, and even educational opportunities to improve aquatic and ecological knowledge.

They also advertise annual events such as beach grass planting and coastal cleanups, which have been incredibly successful in past years. Consider volunteering with DNREC if you’re looking to make an environmental difference!

Delaware Center for the Inland Bays

The Delaware Center for the Inland Bays provides plenty of opportunities to get involved, offering volunteer opportunities that range from science and restoration to education and outreach.

Take part in shoreline maintenance, horseshoe crab monitoring, and more with the Delaware Center for the Inland Bays!

Surfrider Foundation

Volunteer with the Surfrider Foundation’s Delaware Chapter today! The Surfrider Foundation champions many coastal issues, spanning from plastic pollution to ocean protection. One of their most recent accomplishments includes posting signage at Delaware beaches in order to prevent the erosion of planted sand dunes.

To take part in achievements like this, join the Surfrider Foundation and make a difference!

Check It Out!

If you’re interested in learning more about beach ecology from Julia Wingate’s Gallery Interpretation kit, head on over to the Delaware Museum of Nature and Science to see it for yourself.

Education 2025-2026 School Year in Review

As the 2025-2026 school year comes to a close, we want to thank the community for keeping us a part of their experiences.

We interacted with thousands of students across Delaware, Pennsylvania, New Jersey, and Maryland through our educational programming. Thanks to funders like Corteva Agriscience, the Laffey-McHugh Foundation, the Middle Run Foundation, PNC Bank, and more, we were able to do so at little to no cost to many schools.

We welcomed 447 classes for Discovery Tours, 271 of which received partial or full grant funding to cover their programs and transportation. Some of these included attending the third annual Breaking Into STEAM Education event, where middle school students learned about more than 27 careers in our region through hands-on activities.

DelMNS hosted three Delaware Teen Science Cafes as part of the National Teen Science Cafe Network, welcoming 151 teens in grades 7-12 at no cost to meet a STEAM professional, have a free dinner, and explore targeted topics. Some schools elected to offer extra credit to their students for attending, and we thank them for that! The Delaware Teen Science Cafe program was sponsored by Incyte and WSFS.

Work-Based Learning has been a newer focus of our team through career development and exposure opportunities. This year we worked with four high schools: Newark High School, Christiana High School, Glasgow High School, and Odessa High School. Students helped both front-of-house and behind-the-scenes, some with singular tasks, others with a deliverable project by the end of their placement. All experienced memorable moments and learning opportunities as the year progressed. We hope to expand our reach in the next school year as needs arise.

School Day Off Camps were a hit this year! Throughout the year, we hosted 25 day-camps when local districts had the day off, but parents did not. Thank you to the districts that helped spread the word: we are a resource for working families!

We visited 335 classrooms to deliver outreach programs. Of these classrooms, 105 received partial or full grant funding to cover their programs and transportation. This was our third year partnering with schools to offer after-school programming, as well as school-day community partner programming.

Last but certainly not least, we worked closely with 137 educators through our Professional Development workshops; 122 of these seats were fully grant-funded. As requested, our focus this past school year was to develop a Physics-focused theme, and educators had a blast with our staff, learning how to make Physics “phun.”

We look forward to an exciting summer with a plethora of other interactions and especially anticipating a memory-filled 2026-2027 school year starting this Fall.

For more information, please visit our website and feel free to contact our education team at education@delmns.org.

All the Best,
Christina Zampini, B.S.Ed.
Director of Education
(She/Her/Hers)

World Ocean Day

Did you know that oceans cover more than 70% of our planet, yet remain the least explored parts of Earth?

Oceans regulate our climate, support countless ecosystems, and biodiversity. They’re sources of curiosity and wonder; connecting us globally from coast to coast, protecting our oceans means protecting our future!

On World Ocean Day, we spoke with Dr. Liz Shea in the Oceans Hall at the Delaware Museum of Nature and Science. She is the Director of Research and Curator of the Mollusk Collection and cares for and grows the museum’s natural history collection, including the 10th largest mollusk collection in North America.

Liz conducts research on specimens within the collection and brings back new discoveries from the field. She specializes in cephalopod biology, with a special focus on the biodiversity of deep-sea canyons and seamounts. Through her research, Dr. Shea helps uncover the beauty of life found in the ocean’s most unexplored environments.

This video is a collaboration between Delaware Museum of Nature and Science and Reich&Petch (R&P). The Oceans Hall is in the Global Journey Gallery, a part of the museum’s metamorphosis and building renovations completed in 2022 by the Delaware Museum of Nature and Science along with Reich&Petch Design International (Exhibit Designer), The Design Foundation (Media Producer), kubik maltbie (Exhibit Fabricator), JacobsWyper Architects (Architect), Bancroft Construction (General Contractor), and Aegis Property Group (Project Manager).

On Exhibit: Voices in Flight

Bird songs tell rich stories of territory, courtship, and identity.

Explore these stories in three different ways: photographs of birds captured mid-song, scientific sonograms at the museum that visualize each unique vocalization, and audio recordings here.


Each photograph by Lew Scharpf freezes a fleeting moment as a bird lifts its beak and fills the air with sound. The sonograms, on exhibit at the museum and created by Matthew Halley using software by Cornell Lab of Ornithology, translate those songs into a visual language — patterns of frequency and time that make the invisible audible. Many recordings were made by Dr. Halley, with additional audio from xeno-canto.org.

Song Sparrow

(Thyothorus carolinensis)

The pleasing song of this species is a series of trills and clear whistles. It can be heard in pretty much any open or brushy habitat, including residential neighborhoods and farmers’ fields. They live year-round in Delaware and often form small flocks during the breeding season.

Recorded by Matthew R. Halley on 28 April 2020 in Philadelphia Co., PA.

Great Horned Owl

(Thyothorus carolinensis)

The rhythmic, muffled hoots of this species are uttered at very low frequency (less than 1 kHz) — look for the dark lines at the very bottom of the sonogram.

Recorded by David Darrell-Lambert on 17 May 2019 in Essex Co., ON, Canada.

Northern Flicker

(Colaptes auratus)

The loud “fleer” call of this species often alerts birders to its presence in nearby woodlands. Unlike most woodpeckers, flickers are frequently seen foraging on the ground in open areas, eating ants and other insects.

Recorded by Stanislas Wroza on 21 May 2025 in Portneuf, QC, Canada.

Bald Eagle

(Haliaeetus leucocephalus)

The voice of the Bald Eagle is a series of weak chirping whistles — maybe not what you would expect from a
fearsome predator. Because of this, in movies and television programs, the scream of the Red-tailed Hawk (Buteo jamaicensis) is often dubbed in, to make the eagle seem fiercer than it is!

Recorded by Bruce Lagerquist on 16 November 2022 in Clallam Co., WA.

White-breasted Nuthatch

(Sitta carolinensis)

The short nasal calls of this species (“yenk, yenk”) can be heard in open woods with mature trees, especially oaks and pines. They cling to the trunks of trees and can even be seen clambering down head-first, peeking under layers of bark for food.

Recorded by Jim Berry on 21 May 2016 in Chautaqua Co., NY.

Downy Woodpecker

(Dryobates pubescens)

Get to know the squeaky “rattle call” of this species and you will soon be hearing it all over the neighborhood. These small woodpeckers are common residents of our local woodlands and gardens, but more often heard than seen.

Recorded by Jim Berry on 24 August 2016 in Chautauqua Co., NY.

Eastern Phoebe

(Poecile carolinensis)

Listen for the “phoebe!” song and its subtle variants (pictured) in early March. This is one of the first migratory songbirds to return to our region in the spring. Scientists have shown with experiments that the song is encoded in the bird’s DNA — no learning required!

Recorded by Matthew R. Halley on 28 April 2020 in Philadelphia Co., PA.

American Robin

(Turdus migratorius)

The song of this common species is a series of short, warbled phrases. Robins can only hear up to about 8.7 kHZ — the dark horizontal line in the sonogram — which means that we could make a high-pitched whistle that humans could hear, but robins could not!

Recorded by Matthew R. Halley on 29 April 2020 in Philadelphia Co., PA.

Red-tailed Hawk

(Poecile carolinensis)

The raspy scream of this species will send shivers down your spine! These hawks are often seen soaring over fields, tracing lazy circles in the sky as they search for food—small mammals are their preference. They often perch on telephone poles and other man-made structures. But take note—immature red-tailed hawks do not have red tails!

Recorded by Allen T. Chartier on 24 June 2008 in St. Clair Co., MI.

Carolina Wren

(Thyothorus carolinensis)

The song of this species is a rolling series of emphatic whistled phrases, usually 3 or 4 syllables long each. For example, you might hear it singing something that sounds like “tea kettle, tea kettle!” or “generator! generator!” It is commonly heard in woodlots throughout the Mid-Atlantic region.

Recorded by Matthew R. Halley on 22 April 2020 in Philadelphia Co., PA.

Red-bellied Woodpecker

(Melanerpes carolinus)

This common species makes a loud “chhrrrrr” call with a slightly rising inflection. They thrive in open woodlands and often come to backyard suet feeders.

Recorded by Christopher McPherson on 20 May 2019 in Hillsborough Co., NH.

Northern Mockingbird

(Thyothorus carolinensis)

This species is so famous for mimicking the songs of other species that is was given the scientific name “polyglottos,” meaning “speaks many languages.” Sometimes, you will hear mockingbirds singing very late at night. This recording was made after midnight!

Recorded by Matthew R. Halley on 5 June 2022 in Chester Co., PA.

Carolina Chickadee

(Poecile carolinensis)

The song of this familiar species at backyard feeders is a series of four sweet whistles, often referred to by the phrase “fee bee fee bay” or, in this case, “fee bay bay bay.”

Recorded by Thomas Magarian on 13 May 2024 in Page Co., VA.

Pine Siskin

(Poecile carolinensis)

This northern species visits Delaware in the winter and prefers open coniferous forest. Its song is a rapid jumble of husky notes. Look for the “streaky” goldfinch at your bird feeder!

Recorded by Stanislas Wroza on 4 June 2025 in Capitale-Nationale, QC, Canada.

Cedar Waxwing

(Bombycilla cedrorum)

The song of this species is a simple series of high-pitched whistles with an irregular rhythm— “sreeee! sreeee!” The fundamental frequency is usually between 6–8 kHZ. This recording provides a great visual example of Ohm’s Law of acoustics: the first overtone is twice the frequency of the fundamental.

Recorded by Valerie Heemstra on 17 August 2023 in Delta Co., MI.

About the contributors

Lew Scharpf, Ph.D.

Lew Scharpf brings expertise in biochemical research, spanning both industry and academia, to his volunteer work here. He is a musician, photographer and avid birder.

Matthew R. Halley, Ph.D.

Matthew Halley is a scientist, historian, and musician whose work spans evolutionary biology, animal behavior, and the history of science. As the museum’s Research Director and Curator of Birds, he oversees the bird collection and conducts specimen-based research using methods ranging from DNA analysis to audio recording.

A World of Shells

Hidden right in Wilmington, Delaware, the Delaware Museum of Nature and Science houses a true treasure: one of North America’s largest and most important mollusk collections. What started with the curiosity of founder John E. du Pont and legendary malacologist R. Tucker Abbott has grown into a research powerhouse with more than 2.3 million shells from every corner of the globe. From deep-sea cephalopods to bright, delicate land snails, each specimen tells a story of exploration, discovery, and decades of scientific passion.

Thanks to the dedication of curators, researchers, and volunteers, this collection has become an incredible resource for science. Generous donations, global expeditions, and careful cataloging have brought in treasures from the Indo-Pacific, the Gulf of Mexico, South America, and beyond. Digitization projects supported by the National Science Foundation have made thousands of records searchable online, putting Delaware on the map as a key player in global biodiversity research.

What makes this collection truly extraordinary is its connection to people. Museum staff have shared their love of mollusks through exhibits, programs, shell clubs, and partnerships with students and researchers for decades. From the original “Hall of Shells” to today’s Oceans and Delaware ecosystems galleries, visitors get unforgettable views of the beauty and diversity of mollusks.

Mollusk Curators at the Delaware Museum of Nature and Science. 2. R. Tucker Abbott. 3. Rüdiger Bieler. 4. Paula M. Mikkelsen. 5. Timothy A. Pearce. 6. Kevin J. Roe. 7. Elizabeth K. Shea

Read the recent publication in the Nautilus to learn more about the collections.

Career Experiences & Education

In support of our mission to ignite a life-long passion for exploring, discovering, and valuing nature and science, we offer a variety of ways to explore careers within the nature and science fields as they pertain to our museum including Career Awareness, Exploration, and immersive Work-Based Learning.

Career Exploration Opportunities

  • Career Fairs: Students explore a variety of organizations to learn about their careers and what they do. Grades K-12+
  • Guest Speakers: Staff can come to your classroom to discuss multitudes of topics as Subject Matter Experts. Grades K-12+
  • Group Volunteer Opportunities: Looking for a class project? Collaborate to help us deliver our mission in many ways. Grades 6-12+
  • Workplace Tours: See our facility and behind the scenes for all the amazing work we do. Grades 6-12+
  • Mentored Projects: Collaborate with staff for an agreed upon end-goal we can use in our facility. Grades 10-12+
  • Job Shadowing: POV a day in the life of a museum employee. Grades 12+
  • Externships/Practicum: Extended hands-on experience with our staff with targets and goals. Grades 12+

If you are interested in learning more about how your students can participate in Work-Based Learning with us, please email education@delmns.org. 

Work-Based Learning in action

Since 2023, the Delaware Museum of Nature and Science has worked with multiple schools in the Christina School District to provide Work-Based Learning opportunities. Students complete tasks such as helping to feed animals and cleaning supplies, preparing and folding facility materials, stuffing gift bags, stickering library books, greeting guests with maps, counting and cutting craft materials, and more.

New subspecies from the Philippines uncovered in the DelMNS bird collection

Hidden in the drawers of the Delaware Museum of Nature and Science (DelMNS) are treasures with the power to reshape scientific knowledge. In recent articles published in the Journal of Asian Ornithology, Dr. Matthew R. Halley has used the museum’s specimens to describe two new subspecies of “tailorbirds” (songbirds in the genus Orthotomus) and to correct a long-standing error about the plumage of these elusive rainforest birds.


For decades, field guides have depicted males and females of the Black-headed Tailorbird (Orthotomus nigriceps) and White-eared Tailorbird (Orthotomus cinereiceps) as different in color — a pattern that scientists call “sexual dichromatism.” Males are shown with black throats, and females with white or speckled throats. However, after studying a larger sample of specimens than previous scientists, Halley (2022, 2025) has shown that, in the “definitive” (adult) plumage, males and females both have black throats — these species are not “dichromatic” but “monochromatic.” Time to update the field guides!

Photo of White-eared Tailorbird (Orthotomus cinereiceps obscurior) taken on 2 September 2024 by Jay Paul Campaner Mantuon, CC BY-SA 4.0 <https://creativecommons.org/licenses/by-sa/4.0>, via Wikimedia Commons.

Dr. Matthew Halley in the museum’s bird collection, looking at O. c. lucidus study skins.

Halley has also uncovered the existence of two undescribed subspecies, which were overlooked by former ornithologists. First, in 2022, he showed that Black-headed Tailorbird specimens from the islands of Dinagat and Siargao are paler and brighter than those from nearby Mindanao, then named the divergent lineage Orthotomus nigriceps luminosus. Now, in his 2025 paper, Halley presents evidence of a divergent yellow-bellied population of White-eared Tailorbird from the Sarangani Peninsula in south-central Mindanao, which he names Orthotomus cinereiceps lucidus.


These are not just scientific footnotes. Each new subspecies represents an evolutionary lineage that conservationists can now recognize, study, and protect. And each discovery underscores the unique value of the museum’s collections. Without specimens preserved decades ago — carefully cataloged, cared for, and made available to researchers like Halley — these lineages would have remained invisible.

Figure 3 from Halley (2025). Plumage variation in Orthotomus cinereiceps study skins from Mindanao, showing the “grey phenotype” (O. c. obscurior) and “yellow phenotype” (now O. c. lucidus).

Labels attached to the holotype specimens of Orthotomus nigriceps luminosus (top) and O. cinereiceps lucidus (bottom). Type specimens (denoted by red labels) are designated whenever a new species or subspecies is described, to be the official bearers of the proposed scientific name, and to serve as reference for future studies.

References
Halley, M. R. 2022. A new subspecies of Black-headed Tailorbird Orthotomus nigriceps (Cisticolidae) and clarification of age-related plumage sequences. Journal of Asian Ornithology 38: 129–134. LINK

Halley, M. R. 2025. Plumage variation in White-eared Tailorbird Orthotomus cinereiceps (Cisticolidae) with a description of a new subspecies from Mindanao, Philippines. Journal of Asian Ornithology 41: 51–57. LINK

Expedition at Sea

Museum scientist Liz Shea, Ph.D. is at sea on the NOAA Ship Nancy Foster on a research trip. We’ll post updates and reports from Liz here and on our social media!

August 30, 2025: And we’re off!

And we’re off! Over the next 13 days, 10 scientists, three ROV (Remote Operated Vehicle) Global Explorer engineers, and NOAA crew will be at sea together on the NOAA Ship Nancy Foster looking for and sampling deep-sea corals in the Northeast Canyons and Seamounts Marine National Monument. Follow along as we highlight our work each day!

August 31, 2025: Rough Weather

Before the sun set on our first full day, rough weather at the seamounts scrambled our plans. After a discussion with the Commanding Officer, the team set off to find a new spot – Jones Canyon, an unexplored canyon just north of Hudson Canyon. To prepare for our first dive, we stopped to practice deploying and recovering the ROV. The test was successful, and the ROV is ready for operations. We will arrive on site tonight and begin sampling the water column for environmental DNA (eDNA), which helps us to understand what species are lurking beneath the waves and beyond our view. Our first dive will be tomorrow morning – all systems go!

September 1, 2025: Adapt, Adapt, Adapt

At sea, you’ve got to be ready to adapt, adapt, adapt. Before we got to our first station, bad weather offshore caused us to change our original plan, but we adapted and found a new dive site further inshore. When we arrived on site, our water sampling protocol was a challenge at sea, so we adapted again (after 14+ hours of filtering) to streamline our efforts. The ROV was deployed as planned at 0800 this morning, but an unexpected issue brought us to a full stop and recovery of the vehicle. Fair to say it’s been a difficult 24 hours. Happily, our bird survey scientists from U.S. Fish and Wildlife Service found a bright spot for us to focus on – baby dolphins!

September 3, 2025: Setbacks

After a tough first dive, we’ve spent the last 2 days on the dock at the Northeast Fisheries Science Center in Woods Hole, MA. The engineering team is repairing the ROV, the eDNA team is streamlining the workflow to filter water, and the planning team is creating new dive plans to maximize our remaining time at sea. There have been a few frustrating setbacks including the unbelievable coincidence that all our replacement parts for the ROV and additional sampling bags for eDNA were on the same delayed flight out of Louisville, KY. Just another reminder that no matter how much planning is done beforehand, there are no guarantees that everything will go without a hitch. We remain optimistic that we will be headed out to the seamounts soon.

September 5, 2025: Bear or Bust!

The parts came for the ROV, and we left Woods Hole Thursday at 2:30 p.m. with all the gear repaired.  We stopped near Martha’s Vineyard for a dunk test and a few system adjustments, and then got the all clear to head to Bear Seamount. It will take almost 24 hours to get there, but we are excited to get back to exploring! 

September 6, 2025: Exploring Bear Seamount

We arrived at Bear Seamount and the conditions were finally just right for a full ROV dive. The Global Explorer (GEX) was deployed at 8 a.m., and reached bottom two hours later, more than 2000 m beneath the surface. Following the pre-determined waypoints, we surveyed and explored a small part of the eastern side of the seamount. We encountered several different habitat types ranging from steep, rocky slopes, to depressions filled with rubble, coral skeletons, and soft sediments. A few of these even contained skate egg cases! We were excited to see that Bear Seamount is home to such a diverse assemblage of organisms, including hard and soft corals, sponges and a variety of other invertebrates including brittle stars, sea stars, and crinoids (sea lilies).

September 7, 2025: Collecting Specimens

After seven hours exploring Bear Seamount, the ROV was recovered, secured on the back deck, and samples were retrieved from the various collecting containers. During the dive, we collected corals, and other invertebrates including sea stars, brittle stars and crinoids (sea lilies) and any unusual animals we encountered. When an animal is collected, a ‘georeferenced’ record is created so researchers will know the exact location where the organism was found. These data include latitude and longitude, as well as depth and date of collection. In the wet laboratory on the ship, we photograph each specimen and take a tissue sample for genetic barcoding. These specimens are then preserved and will be deposited in the invertebrate collections at the Smithsonian Institution, National Museum of Natural History.  

September 8, 2025: Water Sampling

The weather has been relentless – wherever we go, the winds and waves follow. Although conditions were too rough to deploy the ROV, deploying CTDs to collect water for environmental DNA (eDNA) could still be conducted safely. So, we are maximizing our opportunities to collect eDNA samples. Each CTD deployment requires input from multiple scientists and crew members. The spatial ecologist plots the location of the dive, the Survey crew deploys and retrieves the CTD, the eDNA science team identifies the depth at which water samples are taken, and then spends hours (!) filtering the water once it’s recovered from the bottles. 

eDNA is a relatively new technology that is used to identify organisms from small pieces of DNA shed into the water column. If these small bits can be amplified and identified, then an entire benthic community can be characterized in a single water sample!

September 9, 2025: Cups for Science

A main adaptation deep-sea creatures must face is increasing pressure with depth. But how do you explain the extreme forces these creatures face? Styrofoam cups! Increasing pressure causes the tiny air pockets within Styrofoam cups to compress – the deeper you go the smaller the cup shrinks. To help explain this process, scientists and crew aboard the Nancy Foster used their artistic skills to design cups using ordinary markers. The designs often mark something important or interesting about the cruise, a favorite sea creature, or holiday themes. Luckily, you don’t have to be terribly artistic to participate – even the simplest design is improved by a trip to the seafloor and back. Check out the before and after photos to see how much our cups shrank after a trip to 2,000 meters!

September 10, 2025: One Last Attempt

Thanks to the efforts of NOAA Ship Nancy Foster crew and the ROV team, we rallied for one more dive on our way back to shore. A lot of communication and in-the moment decisions allowed this to happen. We got permission to come in on the second tide of the day instead of the first (buying us 6 hours of potential dive time); due to good weather, we made better time travelling back towards shore than expected (+ 4 hours); we opted out of one last eDNA cast so we could leave our position early (+ 2 hours); and we agreed to start operations 2 hours earlier than normal (+ 2 hours). The ROV will deploy at 0630 at Hydrographer Canyon. We found amazing coral habitat at Hydrographer Canyon last year so we are optimistic. This year, we picked a similar depth on the opposite side of the canyon to increase the likelihood of finding corals.

September 10/11, 2025: Success!

Yesterday’s dive did not disappoint! We were able to spend almost 8 hours on bottom looking at corals and collecting samples. As is true for every dive, we observed some animals, behaviors and amazing habitats that we had not seen before. Once the specimens were back on deck, a tissue sample was taken from each one for genetics work, and then each specimen was photographed before being fixed in preservative for long-term storage at the Smithsonian’s National Museum of Natural History. The imagery and samples collected today provide important information that are critical to better understand the natural resources and ecosystems in and around the Monument and will help us to characterize the biodiversity as well as the type and extent of habitats represented. It is always a great feeling to end a cruise on a high note!

Screen shot of a prickly deep-sea crab, standing on a coral skeleton, covered in zoanthids, that has a brittle star (orange colored) wrapped around its branches.

September 13, 2025: Wrap up

Fieldwork is never easy and this trip was plagued with weather and equipment problems. Twenty-four hour water filtering sessions, and dawn to dusk battles to stay upright while photographing birds in the wind and rain were brutal. Nevertheless, the science and ship crews proved their mettle and worked hard to maximize the amount and types of information gathered during the cruise. It will likely be a long time until we are back working in the Monument. Until then, we will process these samples, identify organisms, analyze water samples for eDNA, and annotate photographs and video footage. All the data we collected will move us closer to a better understanding of the Monument.

With any luck, we will be back out next year (but hopefully in June)!